■ Taxonomic and Host Catalogue of the Tachinidae of America North of Mexico
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Genus Tachina Meigen, 1803

Subgenus NOWICKIA Wachtl, 1894

FABRICIA Latreille, 1829: 510 (junior homonym of Fabricia de Blainville, 1828). Type species: Tachina ferox Panzer, 1809, by fixation of O'Hara & Wood (2004: 325) under Article 70.3.2 of ICZN (1999), misidentified as Musca fera Linnaeus, 1761, in the fixation by monotypy of Latreille (1829) [Palearctic].
FABRICIA Robineau-Desvoidy, 1830: 42 (junior homonym of Fabricia de Blainville, 1828). Type species: Tachina ferox Panzer, 1809 (cited as Musca ferox Panzer, in error), by monotypy [Palearctic].
NOWICKIA Wachtl, 1894: 142. Type species: Echinomyia regalis Rondani, 1859 (= Tachina marklini Zetterstedt, 1838), by original designation [Palearctic].
FABRICIELLA Bezzi, 1906: 49 (nomen novum for Fabricia Robineau-Desvoidy, 1830).
METOPOTACHINA Townsend, 1915b: 21. Type species: Echinomyia palpalis Coquillett, 1902, by original designation.
LARVAEVOROPSIS Townsend, 1916f: 24. Type species: Echinomyia dakotensis Townsend, 1892, by original designation.
ECHINOMYODES Townsend, 1916f: 25. Type species: Echinomyodes piceifrons Townsend, 1916, by original designation.
FABRICIODES Townsend, 1916f: 26. Type species: Fabriciodes montana Townsend, 1916 (= Tachina marklini Zetterstedt, 1838), by original designation.
EULARVAEVORA Townsend, 1916f: 27. Type species: Fabriciella ampliforceps Rowe, 1931, by fixation of O'Hara & Wood (2004: 325) under Article 70.3.2 of ICZN (1999), misidentified as Tachina algens Wiedemann, 1830, in the original designation by Townsend (1916f).

actinosa (Reinhard, 1938).– Oregon, Colorado (S&A, 1965).
– Fabriciella actinosa Reinhard, 1938: 8. Type data: holotype female (CNC). Type locality: USA, Colorado, Alamosa.


acuminata (Tothill, 1924).– California, Colorado, Arizona, New Mexico, Mexico (S&A, 1965), Wyoming, Utah (CNC).
– Fabriciella acuminata Tothill, 1924b: 260. Type data: holotype male (USNM). Type locality: USA, Colorado.


ampliforceps (Rowe, 1931).– Alaska, British Columbia to Nova Scotia, south to California, New Mexico, South Dakota, and New York (S&A, 1965), Northwest Territories, Newfoundland (CNC).
– Fabriciella hispida Tothill, 1924b: 265 (junior secondary homonym of Tachina hispida Robineau-Desvoidy, 1830). Type data: holotype male (CNC). Type locality: Canada, Ontario, Sudbury.
– Fabriciella ampliforceps Rowe, 1931: 673. Type data: holotype male (CNC). Type locality: Canada, Nova Scotia, Mahone Bay.

Hosts from Arnaud (1978: 401), as Nowickia hispida (Tothill)*
Lepidoptera, Noctuidae
Polia purpurissata (Grote)

brevirostris (Tothill, 1924).– British Columbia to California and Colorado (S&A, 1965), New Mexico (CNC).
– Fabriciella brevirostris Tothill, 1924b: 264. Type data: holotype male (CNC). Type locality: Canada, British Columbia, Keremeos.


californimyia Arnaud, 1992.– California (S&A, 1965).
– Fabriciella intermedia Reinhard, 1942a: 27 (junior secondary homonym of Tachina intermedia Zetterstedt, 1844). Type data: holotype male (USNM). Type locality: USA, California, San Jacinto Mountains.
– Tachina californimyia Arnaud, 1992: 167 (nomen novum for intermedia Reinhard, 1942).


canadensis (Tothill, 1924).– Alberta, Colorado, Michigan, Québec (S&A, 1965), Yukon, Northwest Territories, Manitoba, Labrador, New York (CNC).
– Fabriciella canadensis Tothill, 1924b: 264. Type data: holotype male (CNC). Type locality: Canada, Québec, Godbout.


compressa (Tothill, 1924).– Washington, Oregon, California, Colorado (S&A, 1965), Nevada, Arizona (CNC).
– Fabriciella compressa Tothill, 1924b: 262. Type data: holotype male (USNM). Type locality: USA, Colorado, Leadville, 10,000 to 11,000 feet.


cordiforceps (Rowe, 1931).– Washington to Colorado, south to California and New Mexico (S&A, 1965), Mexico (CNC).
– Fabriciella cordiforceps Rowe, 1931: 649. Type data: holotype male (CNC). Type locality: USA, Utah, Pine Valley Mountains, 7000 feet.


dakotensis (Townsend, 1892).– Washington to Manitoba to Nova Scotia, south to Mexico (Chihuahua), South Dakota, Michigan, and Georgia (S&A, 1965), California, Utah, Saskatchewan, Labrador (CNC).
– Echinomyia dakotensis Townsend, 1892a: 94. Type data: syntypes, 3 males (2 male syntypes in SEMK) (mention by Townsend, 1939a: 45, of "Ht male" in SEMK is not regarded as a lectotype fixation because the specimen in question cannot be distinguished from among the 2 male syntypes in SEMK). Type locality: USA, South Dakota, Brookings.
– Larvaevoropsis orientalis Townsend, 1916f: 25 (named for florum of Coquillett, 1897, not Walker, 1849, in part). Type data: holotype male (USNM). Type locality: USA, New Hampshire, Franconia.


egula (Reinhard, 1938).– Oregon to Wyoming, south to California and New Mexico (S&A, 1965).
– Fabriciella egula Reinhard, 1938: 9. Type data: holotype male (CNC). Type locality: USA, Colorado, San Luis Valley.


emarginata (Tothill, 1924).– Alaska, Alberta, Colorado (S&A, 1965).
– Fabriciella emarginata Tothill, 1924b: 260, 269. Type data: lectotype male (USNM), by fixation of Walley, 1933: 168 (mention of "type" is regarded as a lectotype fixation). Type locality: USA, Colorado, Tennessee Pass.


evanida (Reinhard, 1953).– California, Utah, Colorado, Arizona (S&A, 1965).
– Fabriciella evanida Reinhard, 1953a: 53. Type data: holotype male (CNC). Type locality: USA, Colorado, Lake City, 8700 feet.


garretti Arnaud, 1994.– Alaska to Montana, south to Oregon and New Mexico, also Michigan to New Brunswick (S&A, 1965), Yukon, Manitoba, Nova Scotia, Vermont, New Hampshire (CNC).
– Fabriciella pilosa Tothill, 1924b: 263 (junior secondary homonym of Servillia pilosa Robineau-Desvoidy, 1830 and Tachina pilosa Walker, 1853). Type data: holotype male (CNC). Type locality: Canada, British Columbia, Hedley. Tachina garretti Arnaud, 1994: 208 (nomen novum for pilosa Tothill, 1924).

hirtidorsum (Tothill, 1924).
– Fabriciella hirtidorsum Tothill, 1924b: 269. Nomen nudum.


invelata (Reinhard, 1953).– California, Oregon (S&A, 1965), Nevada (CNC).
– Fabriciella invelata Reinhard, 1953a: 51. Type data: holotype male (CAS). Type locality: USA, California, Los Angeles County, Tanbark Flat.


latifacies (Tothill, 1924).– British Columbia to Wyoming, south to Nevada and New Mexico (S&A, 1965).
– Fabriciella latifacies Tothill, 1924b: 262. Type data: holotype male (USNM). Type locality: USA, Nevada, Ormsby County.


latiforceps (Tothill, 1924).– British Columbia, Alberta, Montana, Québec, Maine, also Arizona (S&A, 1965), Yukon, Northwest Territories (CNC).
– Fabriciella latiforceps Tothill, 1924b: 266. Type data: holotype male (CNC). Type locality: Canada, Québec, Godbout.
– Fabriciella argentea Rowe, 1931: 654 (mispelled as argentia in key, p. 647). Type data: holotype male (CNC). Type locality: Canada, Alberta, Banff.


latigena (Tothill, 1924).– British Columbia to Montana, south to California, New Mexico, and Kansas, also Michigan (S&A, 1965).
– Fabriciella latigena Tothill, 1924b: 262. Type data: holotype male (CNC). Type locality: Canada, British Columbia, Lillooet, 3000 feet.


longiunguis (Tothill, 1924).– British Columbia (S&A, 1965).
– Fabriciella longiunguis Tothill, 1924b: 265. Type data: holotype male (CNC). Type locality: Canada, British Columbia, Vernon.


lutzi (Curran, 1925).– Washington to Wyoming, south to California and New Mexico (S&A, 1965).
– Fabriciella lutzi Curran, 1925b: 256. Type data: holotype male (AMNH). Type locality: USA, Colorado, Rifle.


margella (Reinhard, 1942).– New Mexico (S&A, 1965).
– Fabriciella margella Reinhard, 1942a: 28. Type data: holotype male (USNM). Type locality: USA, New Mexico, Koehler.


marklini Zetterstedt, 1838.– Palearctic, Alaska to Labrador and New England, south to Colorado in the Rocky Mountains (S&A, 1965, as montana and nivalis, in part; CNC).
– Tachina marklini Zetterstedt, 1838: 634. Type data: syntypes, 2 males and 2 females (MZLU). Type localities: Various localities in northern Sweden.
– Fabriciodes montana Townsend, 1916f: 26 (named for florum of Coquillett, 1897, not Walker, 1849, in part). Type data: holotype male (USNM). Type locality: USA, New Hampshire, White Mountains.
– Fabriciella nivalis Tothill, 1924b: 264. Type data: holotype male (USNM). Type locality: USA, Alaska, Healey.


nearctica Arnaud, 1992.– British Columbia to Colorado, south to California and Arizona (S&A, 1965).
– Fabriciella spinosa Tothill, 1924b: 263 (junior secondary homonym of Tachina spinosa Zetterstedt, 1838). Type data: holotype male (CNC). Type locality: Canada, British Columbia, Peachland.
– Tachina nearctica Arnaud, 1992: 167 (nomen novum for spinosa Tothill, 1924).


nigella (Reinhard, 1938).– Montana, California, Nevada, Utah (S&A, 1965).
– Fabriciella nigella Reinhard, 1938: 10. Type data: holotype male (CNC). Type locality: USA, Montana, Dillon.


orbitalis (Reinhard, 1942).– California to Wyoming and Nebraska, also Utah (S&A, 1965), Colorado (CNC).
– Fabriciella orbitalis Reinhard, 1942a: 26. Type data: holotype male (CNC). Type locality: USA, Utah, Kane County.


palpalis (Coquillett, 1902).– California, New Mexico (S&A, 1965).
– Echinomyia palpalis Coquillett, 1902b: 120. Type data: holotype male (USNM). Type locality: USA, California, Los Angeles County.


picea (Robineau-Desvoidy, 1830).– Nova Scotia, unrecognized species (S&A, 1965).
– Echinomya picea Robineau-Desvoidy, 1830: 44. Type data: holotype female (BMNH). Type locality: Canada, Nova Scotia.


piceifrons (Townsend, 1916).– Northwest Territories to Nova Scotia, south to California, New Mexico, and Massachusetts, also North Carolina (S&A, 1965).
– Echinomyodes piceifrons Townsend, 1916f: 25 (named for algens of Coquillett, 1897, in part, not Wiedemann, 1830). Type data: holotype female (USNM). Type locality: USA, Vermont.


pictilis (Reinhard, 1942).– Oregon (S&A, 1965).
– Fabriciella pictilis Reinhard, 1942a: 25. Type data: holotype male (USNM). Type locality: USA, Oregon, Steens Mountains, Wildhorse Canyon, 4270-6000 feet.


planiforceps (Tothill, 1924).– North Carolina (S&A, 1965).
– Fabriciella planiforceps Tothill, 1924b: 261. Type data: holotype male (USNM). Type locality: USA, North Carolina.


plumasana (Reinhard, 1953).– California, Arizona (S&A, 1965), Oregon (CNC).
– Fabriciella plumasana Reinhard, 1953a: 53. Type data: holotype male (CAS). Type locality: USA, California, Plumas County, 4 miles west of Quincy.


tahoensis (Reinhard, 1938).– British Columbia, California, Nevada, Arizona (S&A, 1965).
– Fabriciella tahoensis Reinhard, 1938: 11. Type data: holotype female (USNM). Type locality: USA, Nevada, Tahoe Lake.


Subgenus RHACHOGASTER Townsend, 1915


RHACHOGASTER Townsend, 1915d: 291. Type species: Rhachogaster kermodei Townsend, 1915 (= Tachina algens Wiedemann, 1830), by original designation.
UPODEMOCERA Townsend, 1915m: 228 (spelled Eupodermocera by Tothill, 1924b: 258, in error). Type species: Upodemocera robinsoni Townsend, 1915 (= Jurinia nitida van der Wulp, 1882 (preoccupied)), by original designation.


algens Wiedemann, 1830.– Alaska and Northwest Territories to Newfoundland, south to California, Arizona, and Massachusetts (S&A, 1965), New Mexico (CNC).
– Tachina algens Wiedemann, 1830: 285. Type data: lectotype female (NHMW), by fixation of Townsend, 1939a: 60 (mention of "Ht female" is regarded as a lectotype fixation for the female in NHMW bearing Townsend's holotype label). Type locality: North America.
– Echinomya lapilaei Robineau-Desvoidy, 1830: 44. Type data: type(s) (MNHN or lost). Type locality: Canada, Newfoundland.
– Tachina florum Walker, 1849: 722 (junior homonym of Tachina florum Robineau-Desvoidy, 1830). Type data: lectotype male (BMNH), by present designation. Type locality: Canada, Nova Scotia.
– Echinomyia lugubris van der Wulp, 1883: 20. Type data: lectotype male (ISNB), by fixation of Tothill, 1924b: 258 (mention of male "type" is regarded as a lectotype fixation; male specimen labelled as "Type" in ISNB is accepted as lectotype). Type locality: Canada, Québec, Québec City.
– Rhachogaster kermodei Townsend, 1915d: 291. Type data: holotype male (USNM). Type locality: Canada, British Columbia, Penticton.

Hosts from Arnaud (1978: 401), as Nowickia algens (Wiedemann)*
Lepidoptera, Noctuidae
Apamea lignicolor (Guenée), Euxoa messoria (Harris)

eurekana (Reinhard, 1942).– British Columbia, California (S&A, 1965).
– Fabriciella eurekana Reinhard, 1942a: 28. Type data: holotype male (CNC). Type locality: USA, California, Eureka.


latianulum (Tothill, 1924).– British Columbia to South Dakota, south to California and New Mexico, also Ontario, New Brunswick (S&A, 1965), Saskatchewan, Manitoba, Newfoundland (CNC).
– Fabriciella latianulum Tothill, 1924b: 266. Type data: holotype male (CNC). Type locality: Canada, British Columbia, Royal Oak.


oligoria Arnaud, 1992.– Northwest Territories to Idaho, Colorado and Kansas, also Michigan (S&A, 1965), Alberta to Manitoba and Minnesota (CNC).
– Fabriciella latifrons Tothill, 1924b: 260, 269 (junior secondary homonym of Tachina latifrons Meigen, 1824). Type data: lectotype male (CNC), by fixation of Walley, 1933: 168 (mention of "Holotype" is regarded as a lectotype fixation). Type locality: Canada, "Northwest Territories."
– Tachina oligoria Arnaud, 1992: 167 (nomen novum for latifrons Tothill, 1924).

Hosts from Arnaud (1978: 401), as Nowickia latifrons (Tothill)*
Lepidoptera, Noctuidae
Crymodes devastator (Brace)

robinsoni (Townsend, 1915).– British Columbia to California, Arizona and Kansas, also Wyoming, Ontario and Ohio to Massachusetts (S&A, 1965).
– Jurinia nitida van der Wulp, 1882: 82 (junior secondary homonym of Tachina nitida Zetterstedt, 1838). Type data: holotype female (RMNH). Type locality: USA, Arizona.
– Upodemocera robinsoni Townsend, 1915m: 229. Type data: holotype male (USNM). Type locality: USA, Wyoming, Yellowstone Park.


rostrata (Tothill, 1924).– Alaska to Ontario, south to California and New Mexico (S&A, 1965).
– Fabriciella rostrata Tothill, 1924b: 267. Type data: holotype male (CNC). Type locality: Canada, British Columbia, Penticton.


spineiventer (Tothill, 1924).– British Columbia to Montana, south to California (S&A, 1965), Utah (CNC).
– Fabriciella spineiventer Tothill, 1924b: 266. Type data: holotype male (CNC). Type locality: Canada, British Columbia, Mt. McLean, 8000 feet.


Subgenus TACHINA Meigen, 1803


LARVAEVORA Meigen, 1800: 38. Name suppressed by ICZN, 1963: 339.
ECHINODES Meigen, 1800: 38. Name suppressed by ICZN, 1963: 339.
TACHINA Meigen, 1803: 280. Type species: Musca grossa Linnaeus, 1758 (as grossa Fabricius, in error), by subsequent designation of Brauer, 1893: 489 [Palearctic].
ECHINOMYA Latreille, 1805: 377 (also subsequently spelled Echinomyia, unjustified emendation). Type species: Musca grossa Linnaeus, 1758, by subsequent designation of Latreille, 1810: 444 (this type designation, as cited by Herting, 1984: 84, predates the commonly cited designation of Westwood, 1840: 138) [Palearctic].


There are no species assigned to Tachina (Tachina) in America north of Mexico.


Tachina species

Hosts from Arnaud (1978: 480) for "Tachina" spp.*
Coleoptera, Chrysomelidae
Coptocycla sp.
Hemiptera (Homoptera), Cicadidae
Magicicada septendecim (Linnaeus)
Hymenoptera, Apidae
Bombus spp.
Lepidoptera, Aegeriidae
Vespamima pini (Kellicott)
Lepidoptera, Arctiidae
Hyphantria textor (Harris)
Lepidoptera, Geometridae
Cankerworm(-s) (including Alsophila pometaria (Harris) and/or Paleacrita vernata (Peck))
Lepidoptera, Lasiocampidae
Malacosoma americanum (Fabricius)
Lepidoptera, Noctuidae
Alabama argillacea (Hübner), Papaipema nebris (Guenée), Pseudaletia unipuncta (Haworth), Trichoplusia ni (Hübner)
Lepidoptera, Psychidae
Thyridopteryx ephemeraeformis (Haworth)
Lepidoptera, Pyralidae
Acrobasis indigenella (Zeller), Crambus vulgivagellus Clemens
Lepidoptera, Saturniidae
Antheraea polyphemus (Cramer), Callosamia promethea (Drury), Platysamia cecropia (Linnaeus)
Lepidoptera, Sphingidae
Hyles lineata (Fabricius)
Orthoptera, Phasmatidae
Bacillus rossia (Rossi), Diapheromera femorata (Say)

Reference:


Arnaud, P.H., Jr. 1978. A host-parasite catalog of North American Tachinidae (Diptera). United States Department of Agriculture. Miscellaneous Publication 1319: 1–860.


*Host names (family and species) have not been changed from those given in Arnaud (1978). Each host is listed under the appropriate current tachinid name, with the tachinid name used in Arnaud (1978) cited if different from the current one. For more information about Arnaud (1978), and to see a complete list of tachinid names used in that work and their modern equivalents, click here.

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