■ Taxonomic and Host Catalogue of the Tachinidae of America North of Mexico
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Genus Lixophaga Townsend, 1908

LIXOPHAGA Townsend, 1908: 86. Type species: Lixophaga parva Townsend, 1908, by original designation.
EUZENILLIA Townsend, 1911a: 148 (spelled Euzenilla by Townsend, 1912a: 111, in error). Type species: Euzenillia aurea Townsend, 1911 (= Hypostena variabilis Coquillett, 1895), by monotypy.
UROPHYLLOPSIS Townsend, 1916a: 625. Type species: Admontia retiniae Coquillett, 1897, by original designation.
EUZENILLIOPSIS Townsend, 1916h: 76. Type species: Euzenilliopsis diatraeae Townsend, 1916, by original designation.
PHRYNOFRONTINA Townsend, 1919c: 579. Type species: Phrynofrontina convexa Townsend, 1919 (= Sturmia discalis Coquillett, 1902), by original designation.
ERYCIOIDES Curran, 1930b: 103. Type species: Erycioides thoracica Curran, 1930, by original designation.
PROLIXOPHAGA Townsend, 1934: 404. Type species: Lixophaga plumbea Aldrich, 1925, by original designation.
MESSIOMYIA Reinhard, 1955d: 124. Type species: Messiomyia triconis Reinhard, 1955, by original designation.


alberta (Curran, 1925).– Alberta (S&A, 1965), Ontario, Québec (CNC).
– Hypostena alberta Curran, 1925a: 154. Type data: holotype male (CNC). Type locality: Canada, Alberta, Slave Lake.


diatraeae (Townsend, 1916).– Neotropical, Cuba; introduced and established in Florida (S&A, 1965), Texas, Georgia (CNC).
– Euzenilliopsis diatraeae Townsend, 1916h: 76. Type data: holotype male (USNM). Type locality: Cuba (not Louisiana as published). Type host: Diatraea saccharalis (Fabricius), Pyralidae.

Hosts from Arnaud (1978: 342)*
Lepidoptera, Arctiidae
Utetheisa venusta Dalman
Lepidoptera, Hesperiidae
Calpodes ethlius (Stoll)
Lepidoptera, Noctuidae
Spodoptera frugiperda (J.E. Smith)
Lepidoptera, Pyralidae
Diatraea canella Hampson, Diatraea crambidoides (Grote), Diatraea saccharalis (Fabricius), Diatraea spp., Zeadiatraea lineolata (Walker)

discalis (Coquillett, 1902).– Minnesota to Maine, south to Georgia, also Alberta, Utah (S&A, 1965), Yukon, British Columbia, New Brunswick, Nova Scotia (CNC).
– Sturmia discalis Coquillett, 1902b: 114. Type data: holotype female (USNM). Type locality: USA, Wisconsin.
– Phrynofrontina convexa Townsend, 1919c: 580. Type data: holotype male (USNM). Type locality: USA, Maryland, Chain Bridge.
– Nearchus ningulus Reinhard, 1964b: 47. Type data: holotype female (CNC). Type locality: USA, Utah, Beaver.

Hosts from Arnaud (1978: 433), as Phrynofrontina discalis (Coquillett)*
Lepidoptera, Lymantriidae
Euproctis chrysorrhoea (Linneaus)

fasciata Curran, 1930.– Michigan, Ohio, New York (S&A, 1965), New Mexico, Ontario, Québec (CNC), West Virginia, Virginia (WVU).
– Lixophaga fasciata Curran, 1930b: 100. Type data: lectotype male (AMNH), by designation of Arnaud, 1963c: 120. Type locality: USA, New York, Tuxedo, Station for the Study of Insects.

Hosts from Arnaud (1978: 345) for sp. near fasciata Curran*
Lepidoptera, Olethreutidae
Petrova a1bicapitana (Busck)

impatiens (Curran, 1925).– Manitoba (S&A, 1965).
– Hypostena impatiens Curran, 1925a: 154. Type data: holotype male (CNC). Type locality: Canada, Manitoba, Teulon.


jennei Aldrich, 1926.– Arkansas, Louisiana, Florida (S&A, 1965).
– Lixophaga jennei Aldrich, 1926e: 18. Type data: holotype male (USNM). Type locality: USA, Arkansas, Siloam Springs. Type host: Cydia pomonella (Linnaeus) [published as Carpocapsa pomonella], Tortricidae.

Hosts from Arnaud (1978: 346)*
Lepidoptera, Olethreutidae
Laspeyresia pomonella (Linnaeus)

mediocris Aldrich, 1925.– Illinois to New Jersey, south to Mississippi and Florida (S&A, 1965), California (CNC).
– Lixophaga mediocris Aldrich, 1925c: 136. Type data: holotype male (USNM). Type locality: USA, Virginia, East Falls Church. Type host: Rhyacionia frustrana (Comstock), Tortricidae.

Hosts from Arnaud (1978: 346)*
Lepidoptera, Gelechiidae
Frumenta nundinella (Zeller)
Lepidoptera, Limacodidae
Cnidocampa flavescens (Walker)
Lepidoptera, Olethreutidae
Grapholitha molesta (Busck), Laspeyresia caryana (Fitch), Rhyacionia buoliana (Schiffermüller), Rhyacionia frustrana (Comstock), Rhyacionia rigidana (Fernald), Suleima helianthana (Riley)

opaca Reinhard, 1945.– Washington, Alberta, Wyoming, Michigan, Québec (S&A, 1965), Saskatchewan, New Brunswick (CNC), West Virginia (WVU).
– Lixophaga opaca Reinhard, 1945: 30 (junior secondary homonym of Anisia opaca van der Wulp, 1890; not renamed by Wood, 1985: 54, pending a revision of Lixophaga). Type data: holotype male (CNC). Type locality: USA, Washington, Blewett.


orbitalis Aldrich, 1926.– California (S&A, 1965).
– Lixophaga orbitalis Aldrich, 1926e: 17. Type data: holotype male (USNM). Type locality: USA, California, Saticoy. Type host: Cydia pomonella (Linnaeus) [published as Carpocapsa pomonella], Tortricidae.

Hosts from Arnaud (1978: 347)*
Lepidoptera, Olethreutidae
Laspeyresia pomonella (Linnaeus)

parva Townsend, 1908.– Texas (S&A, 1965), Ohio (CNC).
– Lixophaga parva Townsend, 1908: 86. Type data: holotype male (USNM). Type locality: USA, Texas, Dallas. Type host: Lixus scrobicollis Boheman, Curculionidae.

Hosts from Arnaud (1978: 347)*
Coleoptera, Curculionidae
Lixus scrobicollis Boheman
Lepidoptera, Olethreutidae
Grapholitha molesta (Busck)

plumbea Aldrich, 1925.– Utah to Ontario and Connecticut, south to Arizona and Florida (S&A, 1965), British Columbia, California, Wyoming, Manitoba, South Dakota (CNC).
– Lixophaga plumbea Aldrich, 1925c: 134. Type data: holotype male (USNM). Type locality: USA, Virginia, East Falls Church. Type host: Rhyacionia frustrana (Comstock), Tortricidae.

Hosts from Arnaud (1978: 348)*
Lepidoptera, Olethreutidae
Grapholitha molesta (Busck), Rhyacionia frustrana (Comstock)
Lepidoptera, Phaloniidae
Phalonia oenotherana Riley
Lepidoptera, Pyralidae
Pyrausta futilalis (Lederer)
Lepidoptera, Tortricidae
Xenotemna pallorana (Robinson)

retiniae (Coquillett, 1897).– Oregon, California (S&A, 1965).
– Admontia retiniae Coquillett, 1897: 54 (also subsequently spelled retinae, in error). Type data: holotype male (USNM) (type series comprises 1 male and 1 female on same pin, not 2 males as cited in original description). Type locality: USA, California, Alameda. Type host: Rhyacionia sp. [published as Retinia sp.], Tortricidae.

Hosts from Arnaud (1978: 502), as Urophyllopsis retiniae (Coquillett)*
Lepidoptera, Olethreutidae
Rhyacionia pasadenana (Kearfott), Rhyacionia spp., Rhyacionia sp. probably

thoracica (Curran, 1930).– New York (S&A, 1965), Québec (CNC).
– Erycioides thoracica Curran, 1930b: 103. Type data: holotype male (AMNH). Type locality: USA, New York, Tuxedo, Station for the Study of Insects.


triconis (Reinhard, 1955).– Texas (S&A, 1965).
– Messiomyia triconis Reinhard, 1955d: 125. Type data: holotype male (CNC). Type locality: USA, Texas.


unicolor (Smith, 1917).– Montana, New Mexico, New York, Labrador, Maine, New Hampshire, Massachusetts, Connecticut (S&A, 1965), Yukon, Washington, Oregon, British Columbia to Newfoundland and Nova Scotia, Ohio, Vermont (CNC), West Virginia (WVU).
– Pilatea unicolor Smith, 1917b: 137. Type data: lectotype male (MCZ), by designation of Wood, 1985: 103 (though a holotype was designated in the original description, it was apparently not labelled as such in MCZ). Type locality: USA, Maine, Orr's Island.
– Lixophaga nigribasis Curran, 1930b: 100. Type data: holotype male (AMNH). Type locality: USA, New York, Tuxedo, Station for the Study of Insects.


variabilis (Coquillett, 1895).– Manitoba to Maine, south to Texas and Florida, also British Columbia, California (S&A, 1965), Alberta, New Brunswick (CNC).
– Hypostena variabilis Coquillett, 1895c: 57. Type data: lectotype female (USNM), by fixation of Coquillett, 1897: 62 (statement "From the type specimen" is regarded as a lectotype fixation); also by designation of Wood, 1985: 100. Type locality: USA, Illinois, Algonquin.
– Euzenillia aurea Townsend, 1911a: 148. Type data: holotype female (USNM). Type locality: USA, Massachusetts, Melrose Highlands.

Hosts from Arnaud (1978: 351)*
Coleoptera, Bostrichidae
Amphicerus bicaudatus (Say)
Coleoptera, Cerambycidae
Oberea bimaculata (Olivier)
Coleoptera, Curculionidae
Lixus scrobicollis Boheman, Sphenophorus pontederiae Chittenden
Hymenoptera, Tenthredinidae
Unidentified Tenthredinidae
Lepidoptera, Noctuidae
Anomis erosa Hübner, Papaipema cataphracta (Grote), Papaipema nebris (Guenée), Stibadium spumosum Grote, Trichoplusia ni (Hübner)
Lepidoptera, Olthreutidae
Ancylis comptana (Frölich), Epiblema otiosanum (Clemens), Epiblema otiosanum (Clemens) probably, Epiblema scudderiana (Clemens), Epiblema strenuanum (Walker), Eumarozia malachitana (Zeller), Grapholitha molesta (Busck), Laspeyresia caryana (Fitch), Laspeyresia pomonella (Linnaeus), Rhyacionia rigidana (Fernald), Suleima helianthana (Riley)
Lepidoptera, Pyralidae
Desmia funeralis (Hübner), Homoeosoma electellum (Hulst), Loxostege similalis (Guenée), Ostrinia nubilalis (Hübner), Ostrinia penitalis (Grote), Ostrinia spp. (including 0. obliteralis (Walker) or 0. penitalis (Grote))
Hosts from Arnaud (1978: 354) for sp. near variabilis (Coquillett)*
Coleoptera, Curculionidae
Hypera nigrirostris (Fabricius)

Lixophaga species

Hosts from Arnaud (1978: 354) for Lixophaga spp.*
Unidentified Coleoptera
Rhynchophora
Lepidoptera, Olethreutidae
Diatraea impersonatella (Walker), Diatraea saccharalis (Fabricius)
Hosts from Arnaud (1978: 433) for Lixophaga spp., as Phrynofrontina spp.*
Lepidoptera, Olethreutidae
Petrova metallica (Busck)
Lepidoptera, Pyralidae
Dioryctria abietella (Denis & Schiffermüller) probably, Dioryctria amatella (Hulst), Dioryctria clarioralis (Walker)

Reference:


Arnaud, P.H., Jr. 1978. A host-parasite catalog of North American Tachinidae (Diptera). United States Department of Agriculture. Miscellaneous Publication 1319: 1–860.


*Host names (family and species) have not been changed from those given in Arnaud (1978). Each host is listed under the appropriate current tachinid name, with the tachinid name used in Arnaud (1978) cited if different from the current one. For more information about Arnaud (1978), and to see a complete list of tachinid names used in that work and their modern equivalents, click here.

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